Dane Cook (Dusty Crophopper),
Julie Bowen (Lil' Dipper),
Ed Harris (Blade Ranger),
Wes Studi (Windlifter),
Regina King (Dynamite),
Teri Hatcher (Dottie),
Hal Holbrook (Mayday),
Dusty (Dane Cook), the famous racing airplane, learns that his engine is damaged, so he must shift gears and find a new career. He joins an elite corps of firefighting aircraft devoted to protecting historic Piston Peak National Park. When a massive wildfire threatens the park, Dusty -- with the help of his fearless colleagues Blade Ranger, Lil' Dipper (Julie Bowen), Windlifter, Cabbie and the Smokejumpers -- learns what it takes to become a real hero.